sample gobject.pc.in file - What do the "@"s mean?



I don't understand the sample gobject.pc.in file in
http://developer.gnome.org/dotplan/porting/ar01s05.html#configure.in-changes.
What do the "@"s mean in following lines:

prefix= prefix@
exec_prefix= exec_prefix@
libdir= libdir@
includedir= includedir@




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